diff --git a/lisp/org-macro.el b/lisp/org-macro.el
index 8927b3a80..5a4890875 100644
--- a/lisp/org-macro.el
+++ b/lisp/org-macro.el
@@ -191,47 +191,50 @@ as strings, where macro expansion is allowed."
 	   (format "\\`EXPORT_%s\\+?\\'" (regexp-opt keywords)))
 	  record)
       (while (re-search-forward "{{{[-A-Za-z0-9_]" nil t)
-	(let* ((datum (save-match-data (org-element-context)))
-	       (type (org-element-type datum))
-	       (macro
-		(cond
-		 ((eq type 'macro) datum)
-		 ;; In parsed keywords and associated node properties,
-		 ;; force macro recognition.
-		 ((or (and (eq type 'keyword)
-			   (member (org-element-property :key datum) keywords))
-		      (and (eq type 'node-property)
-			   (string-match-p properties-regexp
-					   (org-element-property :key datum))))
-		  (save-excursion
-		    (goto-char (match-beginning 0))
-		    (org-element-macro-parser))))))
-	  (when macro
-	    (let* ((value (org-macro-expand macro templates))
-		   (begin (org-element-property :begin macro))
-		   (signature (list begin
-				    macro
-				    (org-element-property :args macro))))
-	      ;; Avoid circular dependencies by checking if the same
-	      ;; macro with the same arguments is expanded at the same
-	      ;; position twice.
-	      (cond ((member signature record)
-		     (error "Circular macro expansion: %s"
-			    (org-element-property :key macro)))
-		    (value
-		     (push signature record)
-		     (delete-region
-		      begin
-		      ;; Preserve white spaces after the macro.
-		      (progn (goto-char (org-element-property :end macro))
-			     (skip-chars-backward " \t")
-			     (point)))
-		     ;; Leave point before replacement in case of
-		     ;; recursive expansions.
-		     (save-excursion (insert value)))
-		    (finalize
-		     (error "Undefined Org macro: %s; aborting"
-			    (org-element-property :key macro)))))))))))
+	(unless (save-match-data (org-in-commented-heading-p))
+	  (let* ((datum (save-match-data (org-element-context)))
+		 (type (org-element-type datum))
+		 (macro
+		  (cond
+		   ((eq type 'macro) datum)
+		   ;; In parsed keywords and associated node
+		   ;; properties, force macro recognition.
+		   ((or (and (eq type 'keyword)
+			     (member (org-element-property :key datum)
+				     keywords))
+			(and (eq type 'node-property)
+			     (string-match-p properties-regexp
+					     (org-element-property :key
+								   datum))))
+		    (save-excursion
+		      (goto-char (match-beginning 0))
+		      (org-element-macro-parser))))))
+	    (when macro
+	      (let* ((value (org-macro-expand macro templates))
+		     (begin (org-element-property :begin macro))
+		     (signature (list begin
+				      macro
+				      (org-element-property :args macro))))
+		;; Avoid circular dependencies by checking if the same
+		;; macro with the same arguments is expanded at the
+		;; same position twice.
+		(cond ((member signature record)
+		       (error "Circular macro expansion: %s"
+			      (org-element-property :key macro)))
+		      (value
+		       (push signature record)
+		       (delete-region
+			begin
+			;; Preserve white spaces after the macro.
+			(progn (goto-char (org-element-property :end macro))
+			       (skip-chars-backward " \t")
+			       (point)))
+		       ;; Leave point before replacement in case of
+		       ;; recursive expansions.
+		       (save-excursion (insert value)))
+		      (finalize
+		       (error "Undefined Org macro: %s; aborting"
+			      (org-element-property :key macro))))))))))))
 
 (defun org-macro-escape-arguments (&rest args)
   "Build macro's arguments string from ARGS.
diff --git a/testing/lisp/test-org-macro.el b/testing/lisp/test-org-macro.el
index bc493b3d1..b277c097b 100644
--- a/testing/lisp/test-org-macro.el
+++ b/testing/lisp/test-org-macro.el
@@ -117,6 +117,23 @@
       (narrow-to-region (point) (point-max))
       (org-macro-initialize-templates)
       (org-macro-replace-all org-macro-templates)
+      (org-with-wide-buffer (buffer-string)))))
+  ;; Macros in a commented tree are not expanded.
+  (should
+   (string-match-p
+    "{{{macro}}}"
+    (org-test-with-temp-text
+	"#+MACRO: macro expansion\n* COMMENT H\n<point>{{{macro}}}"
+      (org-macro-initialize-templates)
+      (org-macro-replace-all org-macro-templates)
+      (org-with-wide-buffer (buffer-string)))))
+  (should
+   (string-match-p
+    "{{{macro}}}"
+    (org-test-with-temp-text
+	"#+MACRO: macro expansion\n* COMMENT H1\n** H2\n<point>{{{macro}}}"
+      (org-macro-initialize-templates)
+      (org-macro-replace-all org-macro-templates)
       (org-with-wide-buffer (buffer-string))))))
 
 (ert-deftest test-org-macro/escape-arguments ()
diff --git a/testing/lisp/test-ox.el b/testing/lisp/test-ox.el
index b91b80d63..d69660178 100644
--- a/testing/lisp/test-ox.el
+++ b/testing/lisp/test-ox.el
@@ -1351,15 +1351,6 @@ Footnotes[fn:2], foot[fn:test] and [fn:inline:inline footnote]
   ;; Throw an error when a macro definition is missing.
   (should-error
    (org-test-with-temp-text "{{{missing}}}"
-     (org-export-as (org-test-default-backend))))
-  ;; Macros defined in commented subtrees are ignored.
-  (should-error
-   (org-test-with-temp-text
-       "* COMMENT H\n#+MACRO: macro1\n* H2\nvalue\n{{{macro1}}}"
-     (org-export-as (org-test-default-backend))))
-  (should-error
-   (org-test-with-temp-text
-       "* COMMENT H\n** H2\n#+MACRO: macro1\n* H3\nvalue\n{{{macro1}}}"
      (org-export-as (org-test-default-backend)))))
 
 (ert-deftest test-org-export/before-processing-hook ()