diff --git a/testing/lisp/test-ox.el b/testing/lisp/test-ox.el
index 79cf3c9e4..0247db75c 100644
--- a/testing/lisp/test-ox.el
+++ b/testing/lisp/test-ox.el
@@ -1363,6 +1363,119 @@ Footnotes[fn:2], foot[fn:test] and [fn:inline:inline footnote]
      (org-export-expand-include-keyword)
      (eq 3 (org-current-level)))))
 
+(ert-deftest test-org/expand-include/links ()
+  "Test links modifications when including files."
+  ;; Preserve relative plain links.
+  (should
+   (string-prefix-p
+    "file:org-includee-"
+    (let* ((subdir (make-temp-file "org-includee-" t))
+	   (includee (expand-file-name "includee.org" subdir))
+	   (includer (make-temp-file "org-includer-")))
+      (write-region "file:foo.org" nil includee)
+      (write-region (format "#+INCLUDE: %S"
+			    (file-relative-name includee
+						temporary-file-directory))
+		    nil includer)
+      (let ((buffer (find-file-noselect includer t)))
+	(unwind-protect
+	    (with-current-buffer buffer
+	      (org-export-expand-include-keyword)
+	      (org-trim (buffer-string)))
+	  (when (buffer-live-p buffer)
+	    (with-current-buffer buffer (set-buffer-modified-p nil))
+	    (kill-buffer buffer))
+	  (when (file-exists-p subdir) (delete-directory subdir t))
+	  (when (file-exists-p includer) (delete-file includer)))))))
+  ;; Preserve relative angular links.
+  (should
+   (string-prefix-p
+    "<file:org-includee-"
+    (let* ((subdir (make-temp-file "org-includee-" t))
+	   (includee (expand-file-name "includee.org" subdir))
+	   (includer (make-temp-file "org-includer-")))
+      (write-region "<file:foo.org>" nil includee)
+      (write-region (format "#+INCLUDE: %S"
+			    (file-relative-name includee
+						temporary-file-directory))
+		    nil includer)
+      (let ((buffer (find-file-noselect includer t)))
+	(unwind-protect
+	    (with-current-buffer buffer
+	      (org-export-expand-include-keyword)
+	      (org-trim (buffer-string)))
+	  (when (buffer-live-p buffer)
+	    (with-current-buffer buffer (set-buffer-modified-p nil))
+	    (kill-buffer buffer))
+	  (when (file-exists-p subdir) (delete-directory subdir t))
+	  (when (file-exists-p includer) (delete-file includer)))))))
+  ;; Preserve relative bracket links without description.
+  (should
+   (string-prefix-p
+    "[[file:org-includee-"
+    (let* ((subdir (make-temp-file "org-includee-" t))
+	   (includee (expand-file-name "includee.org" subdir))
+	   (includer (make-temp-file "org-includer-")))
+      (write-region "[[file:foo.org]]" nil includee)
+      (write-region (format "#+INCLUDE: %S"
+			    (file-relative-name includee
+						temporary-file-directory))
+		    nil includer)
+      (let ((buffer (find-file-noselect includer t)))
+	(unwind-protect
+	    (with-current-buffer buffer
+	      (org-export-expand-include-keyword)
+	      (org-trim (buffer-string)))
+	  (when (buffer-live-p buffer)
+	    (with-current-buffer buffer (set-buffer-modified-p nil))
+	    (kill-buffer buffer))
+	  (when (file-exists-p subdir) (delete-directory subdir t))
+	  (when (file-exists-p includer) (delete-file includer)))))))
+  ;; Preserve relative bracket links with description.
+  (should
+   (string-prefix-p
+    "[[file:org-includee-"
+    (let* ((subdir (make-temp-file "org-includee-" t))
+	   (includee (expand-file-name "includee.org" subdir))
+	   (includer (make-temp-file "org-includer-")))
+      (write-region "[[file:foo.org][description]]" nil includee)
+      (write-region (format "#+INCLUDE: %S"
+			    (file-relative-name includee
+						temporary-file-directory))
+		    nil includer)
+      (let ((buffer (find-file-noselect includer t)))
+	(unwind-protect
+	    (with-current-buffer buffer
+	      (org-export-expand-include-keyword)
+	      (org-trim (buffer-string)))
+	  (when (buffer-live-p buffer)
+	    (with-current-buffer buffer (set-buffer-modified-p nil))
+	    (kill-buffer buffer))
+	  (when (file-exists-p subdir) (delete-directory subdir t))
+	  (when (file-exists-p includer) (delete-file includer)))))))
+  ;; Preserve absolute links.
+  (should
+   (string=
+    "[[file:/foo/bar.org]]"
+    (let* ((subdir (make-temp-file "org-includee-" t))
+	   (includee (expand-file-name "includee.org" subdir))
+	   (includer (make-temp-file "org-includer-")))
+      (write-region "[[file:/foo/bar.org]]" nil includee)
+      (write-region (format "#+INCLUDE: %S"
+			    (file-relative-name includee
+						temporary-file-directory))
+		    nil includer)
+      (let ((buffer (find-file-noselect includer t)))
+	(unwind-protect
+	    (with-current-buffer buffer
+	      (org-export-expand-include-keyword)
+	      (org-trim (buffer-string)))
+	  (when (buffer-live-p buffer)
+	    (with-current-buffer buffer (set-buffer-modified-p nil))
+	    (kill-buffer buffer))
+	  (when (file-exists-p subdir) (delete-directory subdir t))
+	  (when (file-exists-p includer) (delete-file includer))))))))
+
 (ert-deftest test-org-export/expand-macro ()
   "Test macro expansion in an Org buffer."
   (require 'ox-org)